5
The article you linked (Hamer, et al. 1993 [Open Access PDF]) was cited in Mills (2019) [Open Access PDF] which states,
Work in the 1990s isolated a relationship with the Xq28 region on the X chromosome (5 [Hu, et al. (1995)], 6 [Hamer, et al. 1993]). Subsequent studies found similarity in the sexual orientation of identical twins, with genetics explaining ...
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