Orientation columns in the primary visual cortex are known to have (mostly) simple cells in layers 4 and 6, and (mostly) complex cells in layers 2,3 and 5. Orientation columns spanning an entire range of orientations (0-180 degrees) are “grouped” into ocular dominance columns.
My question: Ocular dominance columns are monocular columns, corresponding either to the contralateral or the ipsilateral eye. Complex cells, however, are binocular. How do these two things go together?